Hi Rob,
I wouldn't rely on your top two pics as true evidence of wartime configurations. The top photo is of the Grizzly that belongs to the Military Technology Foundation in Portola Valley CA(where I volunteer). It started life with the split ring cupola with A.A. machine gun bracket with the vision cupola and pedestal mount being added post-war by the Canadian or Portuguese armies. The second tank is a M4A3(75)W and appears to be crewed by re-enactors. It is certainly post-war as flash suppressors weren't used on the M2 .50 cal during WW II.
It should be noted that the early split ring cupola rotated and that could give the impression that the .50 cal was mounted on the rear of the turret. The picture that Gino posted is an example of that. The cupola was rotated to 180 degrees so the .50 cal faced to the rear. With the .50 cal facing the front, the mount could partially obstruct the commander's view forward view from the hatch mounted cupola periscope.