I can tell you exactly what the cause is. You used Krylon paint. Don't use it! They're garbage. I've had a couple kits ruined because of that garbage paint. Use Rustoleum Primer. I've used it with zero issues. Goes on nice and smooth. What's even better - they're wet/dry sandable.
Krylon paint have been banished from my hobby forever. GARBAGE! GARBAGE! GARBAGE!!!