Sorry folks, that wasn't supposed to be that difficult. And it certainly wasn't a Yank question.
The answer I expected to hear was HMS Redoubt, who towed the lighter that SubLT Culley flew his Sopwith Camel from on August 11, 1918. Culley took 30 minutes to climb up to the required altitude to reach the Zeppelin L53 that had been shadowing the cruisers and destroyers in the North Sea, and he then shot it down. http://www.ww2aircraft.net/forum/album/showphoto.php?photo=14598
Camels were flown from lighters towed by destroyers earlier without achieving kills. The earliest I have found is June 11, 1918, using a Felixstowe F2A flying boat on the lighter, towed behind HMS Teazer.
New question, and back to Yank Surface warfare specialty material (apologies to Mr. Death):
On USS New Jersey's Viet Nam commission (kind of a British term, but I refer to her period of activity during the Viet Nam war), why were the forward 40mm gum tubs painted blue inside?
Rick